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Malkmus, Jonas. Cis-Regulation of Gremlin1 Expression during Mouse Limb Bud Development and its Diversification during Vertebrate Evolution. 2022, Doctoral Thesis, University of Basel, Faculty of Science.

Ramos Martins, Laurène. The Gremlin1 cis-regulatory landscape: a paradigm to study enhancer cooperation in regulation of transcription dynamics. 2020, Doctoral Thesis, University of Basel, Faculty of Science.